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1 National Institute for Agricultural Research, Quantitative and Applied Genetics Unit, 78352 Jouy-en-Josas Cedex, France
This paper presents a pedagogical argument to explain to quantitative geneticists how REML can be derived from maximum likelihood for estimation of variance components. The argument is first developed for N independent normal observations with mean µ and variance
and is afterward extended to a general linear mixed model structure, y
N(Xß,V). The argument is taken from expectation-maximization theory and consists of replacement of a quadratic in µ or ß by its conditional expectation given the observed data and the variance components.
Key Words: variance components mixed model restricted maximum likelihood expectation-maximization
Submitted on November 9, 1992
Accepted on March 19, 1993
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